Shinra Kusakabe carries the weight of a "devil" and the power of a third-generation pyrokinetic in the Solar Year 198, where Tokyo is plagued by spontaneous human combustion. As the newest recruit of Special Fire Force Company 8, he rockets into battle leaving scorched "devil’s footprints" behind destroying his shoes in the process to save citizens from turning into mindless Infernals. He finds a vital anchor in Sister Iris, a nun whose serene prayers offer a spiritual sanctuary against the destructive fire burning within him. His quest for heroism takes a dark turn when a mysterious cult reveals that the fire which claimed his family was the spark of a global conspiracy. Now, Shinra must navigate dangerous secrets to prevent the world from being consumed by flames. He is determined to uncover the truth and prove that a boy with the footprints of a demon can truly be a hero.